September 26, 2014
Ray D. Owen, who died on Sept. 21 in Pasadena, California, at the age of 98, discovered the phenomenon of immune tolerance, fueling a revolution in immunology and laying the foundation for the successful transplantation of human organs. Owen left Madison in 1947 to join the faculty at Caltech, where he remained for the rest of his long, distinguished career. His later work included studies on human antibodies, blood-group antigens, and the evolution of immune systems.